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In Nazi occupied France, young Jewish refugee Shosanna Dreyfus witnesses the slaughter of her family by Colonel Hans Landa. Narrowly escaping with her life, she plots her revenge several years later when German war hero Fredrick Zoller takes a rapid interest in her and arranges an illustrious movie premiere at the theater she now runs. With the promise of every major Nazi officer in attendance, the event catches the attention of the "Basterds", a group of Jewish-American guerilla soldiers led by the ruthless Lt. Aldo Raine. As the relentless executioners advance and the conspiring young girl's plans are set in motion, their paths will cross for a fateful evening that will shake the very annals of history.

During World War II, Lt. Aldo Raine leads his squad of Jewish-American soldiers behind enemy lines in Nazi occupied France. Their job is simple: kill the enemy. They also have a particularly violent approach to what they do scalping their victims. Soon they are known and feared throughout the German army. In Paris, Shosanna runs a cinema and through a chance meeting with a German war hero, is selected to host the premiere of one of Dr. Josef Goebbels propaganda film. With all of the German high command scheduled to attend - including Hitler, Goering and Martin Bormann - it seems like the ideal opportunity for Lt. Raine and his men to bring the war to a quick end. Shosanna, who is Jewish and whose parents and siblings were killed before her eyes, also has her own plans for the evening's festivities. When she realizes that the man who killed her family Colonel Hans Landa, known as the Jew Hunter, will also be there it makes her own plans complete.

During World War II some American soldiers are sent into Europe to wipe out as many Germans as they can. They've been so good at it that they become known as The Basterds. They eventually learn of an opportunity to get as many high ranking Germans including during the screening of a movie about a heroic German soldier. But things change when all of the members of the team who can speak German are killed and when the German whose the subject of the movie is attracted to the owner of a cinema. He has the screening moved there. What he doesn't know is that the owner is a Jew who a few years ago saw her family killed by German officer who was tasked with rounding all the Jews. She decides to kill them herself.

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